Can once in a lifetime happen twice?
When Jonathan and Sara meet for the first time, there's a definite attraction between the two of them, but they're both attached. Sara is a big believer in "meant to be", and when she gives Jonathan her phone number only to have the wind blow the number from his hand, she sees this as a sign that the two of them are not meant to be together - at least not right now.
Years pass, and now Jonathan (John Cusack) is engaged to marry someone else. He loves his fiancee, but suddenly he's thinking a lot about Sara. Across the country, Sara (Kate Beckinsale) has also just gotten engaged, but she's starting to wonder about Jonathan. They're both starting to wonder what might have been.
The two launch a search to find one another, armed with very little information about the other - they don't even know each other's last name, let alone an address or phone number.